Known as a bully, Frodo the chimpanzee was Gombe's alpha male for five years. (IMAGE)

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Known as a bully, Frodo the chimpanzee was Gombe's alpha male for five years.

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Known as a bully, Frodo the chimpanzee was Gombe's alpha male for five years.

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Ian C. Gilby, Arizona State University.

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Credit Needed: Ian C. Gilby, Arizona State University.

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